- Foreword to the French Edition
- Foreword to the English Edition
- Preface
- 1. What We Know about the Nature of the Horse
- Then and Now
- Discovering the Real Life of Free-Ranging Horses
- Equine Ethology Studies to Pursue
- The Emergence of a New Field of Research: The Cognitive Ethology of the Horse
- 2. Equine Intelligence
- Are Horses Smart? One Question, Several Answers
- A Rash of Clever Horses
- 3. Animal Intelligence, Cognition, and Representation
- Intelligence and Cognition
- Animal Behavior, Cognition, and Representation
- 4. The Equine Brain
- Nervous Tissue and the General Organization of the Mammalian Nervous System
- Brain and Mind in the Light of Evolution
- 5. The Nature of Equine Perception
- Perception: A Dynamic Process That Constructs the World
- A Few Issues Regarding the Study of Equine Perception
- 6. The Anatomical and Physiological Basis of Equine Visual Perception
- Size, Arrangement of the Eyes, and Visual Field
- Anatomical Structure of the Eye
- A Short Tour of the Anatomy of the Retina
- Structure of the Retina and Visual Quality
- Optical Pathways and Cortical Distribution
- Chromatic Theory and Color Perception
- 7. The Behavioral Exploration of Equine Visual Perception: Perception of Shapes and Movement
- Experimental Procedures
- Visual Acuity
- The Visual Field
- Night Vision
- The Visual Apparatus: An Integrated System…
- …In the Context of Cerebral Hemispheric Specialization
- Perceiving the Third Dimension
- Image Recognition
- Object Recognition
- Perceiving Movement
- The Equine Visual Environment: Seen as a Whole or the Sum of Its Parts?
- 8. The Behavioral Exploration of Equine Visual Perception: The Quest for Color Perception
- Brightness: A Vexing Dimension
- A Pioneering Study (Grzimek 1952)
- An Inconclusive Replication (Pick et al. 1994)
- An Apparent Confirmation of Grzimek’s Results (Smith and Goldman 1999)
- New Uncertainties Centering on Brightness (Macuda and Timney 1999)
- The Evidence for a Neutral Point (Geisbauer et al. 2004)
- Color Preferences (Hall et al. 2005)
- Do Horses Perceive the Entire Color Spectrum? (Hall et al. 2006)
- The Neutral Point: Break or Continuity? (Roth, Balkenius, and Kelber 2007)
- Equine Dichromacy: A Qualification (Hanggi, Ingersoll, and Waggoner 2007)
- A New Experiment in Chromatic Discrimination (Blackmore et al. 2008)
- How Well Do Horses Discriminate Color in Half-Light? (Roth, Balkenius, and Kelber 2008)
- Colors That Can Be Fairly Well Discriminated across the Light Spectrum (Timney and Macuda 2009)
- A Provisional Summing Up
- 9. Hearing in Horses
- Nature, Representation, and Characterization of Acoustic Information
- The Equine Auditory System: Anatomy and Physiology
- Behavioral Exploration of Equine Auditory Perception
- 10. Equine Chemical Perception: Odors, Pheromones, Tastes, and Flavors
- Olfactory Perception in the Horse
- From Taste to Flavor
- 11. Tactile Perception in the Horse
- Structure and Function of Horse Skin
- Receptors: Equine Sensory Pathways and Skin Sensitivity
- Mutual Grooming and Neurophysiological Response
- Tactile Stimulation and Interspecific Social Relationships
- Conclusion
- References
- Acknowledgments
- Index
[Color plates follow page 218]





